Order: PHACOPIDA Salter, 1864
Suborder: PHACOPINA Struve, 1959
Superfamily: ACASTOIDEA Reijers, 1972
Family: ACASTIDAE Delo, 1935
Subfamily: ASTEROPYGINAE Delo, 1935
Genus DESTOMBESINA Morzadec, 1997
Diagnosis
by Bignon and Crônier, 2014
Cephalon: Frontal lobe diamond shaped; axial furrows straight between S1 and S3; L3 above L1 and L2; anteriormost portion of the palpebral lobe posterior to intersection of S3 and axial furrow, 4 lenses per dorsoventral file of visual surface; genal spines weak, shorter than glabella with a narrow proximal portion; lateral border rather large, undeveloped on genal spine.
Pygidium: Pygidium with anterior pleural bands lower and narrower than posterior, flat pleural bands; axis with 6–7 rings, constricted between the 3rd and 4th rings; 4–5 pleural segments; 4 pairs of reduced pygidial spines, mainly developed from posterior pleural bands; terminal pygidial spine larger than axis, triangular, twice as long as other pygidial spines.
Occurence
There are two known species who both occur in the upper Emsian. But only Destombesina tafilaltensis is found in Morocco.
Species
Destombesina tafilaltensis, Morzadec 2001 (Upper Emsian in Morocco)
Destombesina ougartensis, Morzadec 1997 (Upper Emsian in Algeria)
References
Bignon, A., Crônier, C., 2014. The systematics and phylogeny of the Devonian subfamily Asteropyginae (Trilobita: Phacopida). Journal of Systematic Palaeontology 12, 637–668.